No caeca; oil-gland tufted; 15 cervical vertebrae. Sternum with four notches on the posterior margin; manubrium simple, composed of the spina externa only. Both carotids are present. Spinal feather-tract well denned on the neck and not forked on the back, so there is no dorsal apterium. Down present in adults on the tracts not covered by feathers, a character absent in other suborders of Anisodactyli. No accessory semitendinosus muscle. The two plantar tendons are generally united, but the slip leading to the hallux branches off from the flexor perforans digitorum above the junction as in Meropes. In some cases however, as in Dacelo, this slip unites with the flexor longus hallucis, and the latter runs to the fourth digit only and has no other connexion with the f. perf. digitorum (Gadow in Newton's Dictionary of Birds, p. 617).